William Karasov

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Professor

226 Russell Labs
1630 Linden Drive
Madison, WI 53706
Phone: 608/263-9319 (office)
608/263-4344 (lab)

Email: wkarasov@wisc.edu

Lab Website

Expertise

Physiological ecology of terrestrial vertebrates.

Education/Experience

Education

Degree Institution Major Field Granted
B.A. University of Minnesota Biology 1975
M.S. U.C.L.A. Biology 1981
Ph.D. U.C.L.A. Biology 1984

Research

Professor Karasov is interested in the physiological ecology of terrestrial vertebrates, particularly the ecological implications of how they process energy, nutrients, and toxins.

Nutritional ecology

The relationships between the nutrient requirements of animals and the nutritional value of their food resources potentially affect diet selection, productivity, and survival. Professor Karasov and his students have performed laboratory and field studies on the nutritional ecology of several species of mammals and birds in order to explore the significance of nutrition in their ecology. This work includes studies of how antiherbivory chemicals in plants affect food selection, digestion, physiology, and population biology of herbivores.

Digestive ecology

Vertebrates differ considerably in the types of foods that they consume, but little is known of the physiological adaptations required for effective utilization of alternate food types. Professor Karasov and his students are studying the digestive physiology of animals with different dietary habits (such as carnivores, herbivores, omnivores, frugivores, and nectarivores) and also how digestive features vary over the short-term (i.e., within a species) and between species (i.e., over evolutionary time).

Ecological energetics

All organisms need energy, and many activities of animals are centered around how to get, process it, and conserve it. Professor Karasov and his students are studying the energetics of free-living animals using doubly labeled water, micrometeorology, telemetry, and direct observation of behavior. They are documenting the physical, physiological, and behavioral factors which affect energy intake and expenditure, and the relationships between foraging behavior and reproductive success.

Wildlife Toxicology

Animals living in or feeding from the Great Lakes are exposed to a number of pollutants, notably halogenated organic compounds and heavy metals. Professor Karasov and his students are participating in studies to determine the level of exposure of fish-eating birds and amphibians, and whether the exposure causes physiological and behavioral dysfunction and effects on population biology.

International and Other Activities

Professor Karasov has served as graduate advisor for students from around the world. Additionally, he has lectured, taught and done research in a variety of locations such as: Argentina, China, Israel, Taiwan, United Kingdom, and South Africa.

Professor Karasov has served on the editorial boards of several journals: Functional Ecology (2002-2004), The Auk (2000-2004), Physiological and Biochemical Zoology (1998-2001), Ecology (1995-1997), and American Journal of Physiology (1993-2001).

Courses Taught

FWE 401 Physiological Animal Ecology – Physiological adaptation and function in wild animals, primarily birds, mammals, reptiles, amphibians. Focus on interactions between animals and their environment, and relationships between animal physiology and the ecology and dynamics of populations. 

Physiological Animal Ecology

 

Ecotoxicology

Publications

Selected Publications

Google Scholar Profile.

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+Journals - 227

Anderson, R. A., and W. H. Karasov (1981) Contrasts in energy intake and expenditure in sit‑and‑wait and widely foraging lizards.  Oecologia 49:67‑72.

Karasov, W. H. (1981) Daily energy expenditure and the cost of activity in a free‑living mammal.  Oecologia 51:253‑259.

Karasov, W. H. (1982) Energy assimilation, nitrogen requirement, and diet in free‑living antelope ground squirrels, Ammospermophilus leucurus.  Physiological Zoology 55:378‑392.

Karasov, W. H. (1983) Water flux and water requirement in free‑living antelope ground squirrels, Ammospermophilus leucurus.  Physiological Zoology 56:94‑105.

Karasov, W. H. (1983) Wintertime energy conservation by huddling in antelope ground squirrels, Ammospermophilus leucurus.  Journal of Mammalogy 64:341‑345.

Karasov, W. H., and J. M. Diamond (1983) A simple method for measuring intestinal solute uptake in vitro.  Journal of Comparative Physiology 152:105‑116.

Karasov, W. H., and J. M. Diamond (1983) Adaptive regulation of sugar and amino acid transport by vertebrate intestine.  American Journal of Physiology 245:G443‑462.

Karasov, W. H., R. S. Pond III, D. H. Solberg, and J. M. Diamond (1983) Regulation of proline and glucose transport in mouse intestine by dietary substrate levels.  Proceedings of the National Academy of Science, USA 80:7674‑7677.

Karasov, W. H., and R. A. Anderson (1984) Interhabitat differences in energy acquisition and expenditure in a lizard.  Ecology 65:235‑247.

Diamond, J. M., and W. H. Karasov (1984) Effect of dietary carbohydrate on monosaccharide uptake by mouse small intestine in vitro.  Journal of Physiology 349:419‑440.

Karasov, W. H. (1985) Nutrient constraints in the feeding ecology of an omnivore in a seasonal environment.  Oecologia 66:280‑290.

Karasov, W. H., and J. M. Diamond (1985) Digestive adaptations for fueling the cost of endothermy.  Science 228:202‑204.

Karasov, W. H., D. H. Solberg, and J. M. Diamond (1985) What transport adaptations enable mammals to absorb sugars and amino acids faster than reptiles?  American Journal of Physiology 249:G271‑283.

Karasov, W. H., D. H. Solberg, S. D. Chang, E. D. Stein, M. Hughes, and J. M. Diamond (1985) Is intestinal transport of sugars and amino acids subject to critical‑period programming?  American Journal of Physiology 249:G770‑785.

Diamond, J. M., W. H. Karasov, D. Phan, and F. L. Carpenter (1986) Hummingbird digestive physiology, a determinant of foraging bout frequency.  Nature 320:62‑63.

Karasov, W. H., E. Petrossian, L. Rosenberg, and J. M. Diamond (1986) How do food passage rate and assimilation differ between herbivorous lizards and nonruminant mammals?  Journal of Comparative Physiology 156:599‑609.

Karasov, W. H., D. Phan, J. M. Diamond, and F. L. Carpenter (1986) Food passage and intestinal nutrient absorption in hummingbirds.  Auk 103:453‑464.

Karasov, W. H., D. Solberg, S. Carter, M. Hughes, D. Phan, F. Zollman, and J. M. Diamond (1986) Uptake pathways for amino acids in mouse intestine.  American Journal of Physiology 251:G501-G508.

Karasov, W. H. (1986) Energetics, physiology, and vertebrate ecology.  Trends in Ecology and Evolution 1:101‑104.

Karasov, W. H., D. H. Solberg, and J. M. Diamond (1987) Dependence of intestinal amino acid uptake on dietary protein or amino acid levels.  American Journal of Physiology 252:G614‑G625.

Karasov, W. H., and E. S. Debnam (1987) Rapid adaptation of intestinal glucose transport: a brush‑border or basolateral phenomenon?  American Journal of Physiology 253:G54-61.

Diamond, J. M., and W. H. Karasov (1987) Adaptive regulation of intestinal nutrient transporters.  Proceedings of the National Academy of Science 84:2242-2245.  USA.

Anderson, R. A., and W. H. Karasov (1988) Energetics of reproduction in a wide foraging lizard, Cnemidophorus tigris, and life history consequences of food acquisition mode.  Ecological Monographs 58:79-110.

Karasov, W. H., L. R. Han, and J. C. Munger (1988) Measurement of D2O by infrared absorbance in doubly labeled water studies of energy expenditure.  American Journal of Physiology 255:R174-177.

Debnam, E. S., W. H. Karasov, and C. S. Thompson (1988) Nutrient uptake by rat enterocytes during diabetes mellitus: Evidence for an increased sodium electrochemical gradient.  Journal of Physiology (London) 397:503-512.

Karasov, W. H., and J. M. Diamond (1988) Interplay between physiology and ecology in digestion.  BioScience 38(9):602-611.

Martinez del Rio, C., W. H. Karasov, and D. J. Levey (1989) Physiological basis and ecological consequences of sugar preferences in cedar waxwings.  Auk 106:64-71.

Munger, J. C., and W. H. Karasov (1989) Sublethal parasites and host energy budgets:  Tapeworm infection in white-footed mice.  Ecology 70:904-921.

Meyer, M., and W. H. Karasov (1989) Antiherbivore chemistry of Larrea tridentata:  Effects on desert woodrat (Neotoma lepida) feeding and nutrition.  Ecology 70:953-961.

Levey, D. J., and W. H. Karasov (1989) Digestive responses of temperate birds switching to fruit and insect diets.  Auk 106:675-686.

Munger, J. C., W. H. Karasov, and D. Chang (1989) Host genetics as a cause of overdispersion of parasites among hosts:  How general a phenomenon?  Journal of Parasitology 75(5):707-710.

Karasov, W. H. (1989) Nutritional bottleneck in an herbivore, the Desert Woodrat (Neotoma lepida).  Physiological Zoology 62(6):1351-1382.

Martinez del Rio, C., and W. H. Karasov (1990) Digestion strategies in nectar- and fruit-eating birds and the sugar composition of plant rewards.  American Naturalist 136:618-637.

Karasov, W. H., and D. J. Levey (1990) Digestive system tradeoffs and adaptations of frugivorous birds.  Physiological Zoology 63:1248-1270.

Munger, J. C., and W. H. Karasov (1991) Sublethal parasites in White-footed Mice:  Impact on survival and reproduction.  Canadian Journal of Zoology 69:398-404.

Karasov, W. H., B. W. Darken, and M. C. Bottum (1991) Dietary regulation of intestinal ascorbate uptake in guinea pigs.  American Journal of Physiology 260:G108-118.

Titus, E., W. H. Karasov, and G. A. Ahearn (1991) Dietary modulation of intestinal nutrient transport in the teleost fish tilapia.  American Journal of Physiology 261:R1568-1574.

Karasov, W. H., M. C. Brittingham, and S. A. Temple (1992) Daily energy expenditure by Black-capped Chickadees (Parus atricapillus) during winter.  Auk 109:393-394.

Karasov, W. H. (1992) Daily energy expenditure and the cost of activity in mammals.  American Zoologist 32:238-248.

Dykstra, C. J., and W. H. Karasov (1992) Changes in House Wren gut structure and function in response to increased energy demands.  Physiological Zoology 65:422-442.

Karasov, W. H., M. W. Meyer, and B. W. Darken (1992) Tannic acid inhibition of amino acid and sugar absorption by mouse and vole intestine:  Tests following acute and subchronic exposure.  Journal of Chemical Ecology 18:719-736.

Levey, D. J., and W. H. Karasov (1992) Digestive modulation in a seasonal frugivore, the American Robin (Turdus migratorius).  American Journal of Physiology 262:G711-G718.

Karasov, W. H. (1992) Tests of the adaptive modulation hypothesis for dietary control of intestinal nutrient transport.  American Journal of Physiology 263:R496-502.

Ellner, L. R., and W. H. Karasov (1993) Latitudinal variation in the thermal biology of ornate box turtles.  Copeia 1993(2):447-455.

Nosek, J. A., S. R. Craven, W. H. Karasov, and R. E. Peterson (1993) 2,3,7,8-Tetrachlorodibenzo-p-dioxin in terrestrial environments:  Implications for resource management.  Wildlife Society Bulletin 21:179-187.

Dykstra, C. R., and W. H. Karasov (1993) Nesting energetics of House Wrens (Troglodytes aedon) in relation to maximal rates of energy flow.  Auk 110:481-491.

Hume, I. D., W. H. Karasov, and B. W. Darken (1993) Acetate, butyrate, glucose and proline uptake in the caecum and colon of prairie voles (Microtus ochrogaster).  Journal of Experimental Biology 176:285-297.

Guglielmo, C. G., and W. H. Karasov (1993) Endogenous losses of mass and energy in Ruffed Grouse.  Auk 110:386-390.

Jakubas, W. J., W. H. Karasov, and C. G. Guglielmo (1993) Coniferyl benzoate in quaking aspen:  Its effect on energy and nitrogen digestion and retention in ruffed grouse.  Physiological Zoology 66:580-601.

Jakubas, W. J., W. H. Karasov, and C. G. Guglielmo (1993) Ruffed grouse tolerance and biotransformation of the plant secondary metabolite coniferyl benzoate.  Condor 95:625‑640.

Jakubas, W. J., B. C. Wentworth, and W. H. Karasov (1993) Physiological and behavioral effects of coniferyl benzoate on avian reproduction.  Journal of Chemical Ecology 19:2353‑2377.

Karasov, W. H. (1993) In the belly of the bird.  Natural History 102:32-37.

Dykstra, C. R., and W. H. Karasov (1993)  Daily energy expenditure by nestling house wrens (Troglodytes aedon).  Condor 95:1028-1030.

Munger, J. C., and W. H. Karasov (1994) Costs of bot fly infection in white-footed mice:  Energy and mass flow.  Canadian Journal of Zoology 72:166-173.

Karasov, W. H., and S. J. Cork (1994) Glucose absorption by a nectarivorous bird:  The passive pathway is paramount.  American Journal of Physiology 267:G18-G26.

Levey, D. J., and W. H. Karasov (1994) Gut passage of insects by European starlings and a comparison with other species.  The Auk 111:478-481.

Guglielmo, C. G., and W. H. Karasov (1995) Nutritional quality of winter browse for Ruffed Grouse.  Journal of Wildlife Management 59:427-436.

Afik, D. and W. H. Karasov (1995) Tradeoffs between digestion rate and efficiency in Yellow-rumped Warblers, and their ecological and evolutionary implications. Ecology 76:2247-2257.

Afik, D., E. Caviedes-Vidal, C. Martinez del Rio, and W. H. Karasov (1995) Dietary modulation of intestinal hydrolytic enzymes in Yellow-rumped Warblers.  American Journal of Physiology 269:R413-R420.

Jakubas, W. J., C. G. Guglielmo, C. Vispo, and W. H. Karasov (1995).  Sodium balance in ruffed grouse as influenced by sodium levels and plant secondary metabolites in quaking aspen.  Canadian Journal of Zoology 73:1106-1114.

Karasov, W. H., and S. J. Cork (1996) Test of a reactor-based digestion optimization model for nectar-eating Rainbow Lorikeets.  Physiological Zoology 69:117-138.

Karasov, W. H., D. Afik, and B. W. Darken (1996) Do Northern Bobwhite quail modulate intestinal nutrient absorption in response to dietary change?  A test of an adaptational hypothesis.  Comparative Biochemistry and Physiology:113A:233-238.

Caviedes-Vidal, E., and W. H. Karasov (1996).  Glucose and amino acid absorption in House Sparrow intestine and its dietary modulation.  American Journal of Physiology 271:R561-R568.

Covell, D. F., D. S. Miller, and W. H. Karasov. (1996)  Cost of locomotion and daily energy expenditure by free-living swift foxes (Vulpes velox):  A seasonal comparison.  Canadian Journal of Zoology 74:283-290.

Larson, J. M., J. P. Giesy, B. Handbridge, W. H. Karasov, L. Sileo, and K. Stromborg (1996).  Reproductive success, developmental anomalies, and environmental contaminants in double‑crested cormorants (Phalacrocorax auritus).  Environmental Toxicology  and Chemistry 15:553-559.

Guglielmo, C., W. H. Karasov, and W. J. Jakubas (1996).  Nutritional costs of a plant secondary metabolite explain selective foraging by ruffed grouse.  Ecology 77:1103-1115.

Dykstra, C. R., M. W. Meyer, and W. H. Karasov (1997).  Validation of the doubly-labeled-water method in bald eagles and a comparison of two equations for the calculation of energy expenditure.  Physiological Zoology 70:19-26.

Afik, D., B. W. Darken, and W. H. Karasov (1997) Is diet-shifting facilitated by modulation of intestinal nutrient uptake?  Test of an adaptational hypothesis in yellow-rumped warblers.  Physiological Zoology 70:213-221.

Afik, D., S. R. McWilliams, and W. H. Karasov (1997)  Test for passive absorption of glucose in yellow‑rumped warblers, and its ecological implications.  Physiological Zoology 70:370-377.

Weiser, J. I., A. Porth, D. Mertens, and W. H. Karasov (1997)  Digestion of chitin by Northern Bobwhites (Colinus virginianus) and American Robins (Turdus migratorius).  The Condor 99:553-556.

Karasov, W. H. and R. A. Anderson (1998) Correlates of variation in average daily metabolism of field-active zebra-tailed lizards (Callisaurus draconoides).  Physiological Zoology 71:93-105.

Dykstra, C. R., M. W. Meyer, D. K. Warnke, W. H. Karasov, D. E. Anderson, W. W. Bowerman, IV, and J. P. Giesy (1998).  Low reproductive rates of Lake Superior Bald Eagles: low food delivery rates or environmental contaminants?   Journal of Great Lakes Research 24:32‑44.

Woodford, J.E., W.H. Karasov, M.W. Meyer and L. Chambers (1998) Impact of 2,3,7,8-TCDD exposure on survival, growth, and behavior of ospreys breeding in Wisconsin, USA.  Environmental Toxicology and Chemistry 17:1323-1331.

Huang, Y.W., M.J. Melancon, R.E. Jung, and W.H. Karasov (1998) Induction of cytochrome P450-associated monooxygenases in northern leopard frogs, Rana pipiens, by 3,3’4,4′,5-pentachlorobiphenyl (PCB 126).  Environmental Toxicology and Chemistry 17:1564-1569.

Karasov, W.H. and B. Pinshow (1998) Changes in lean mass and in organs of nutrient assimilation in a long-distance migrant at a springtime stopover site. Physiological Zoology 71(4):435-448.

McWilliams, S. R. and W. H. Karasov (1998) Do variable-reward feeding schedules affect digestive performance of migratory birds?  Oecologia 114:160-169.

Dearing, M.D., A.M. Mangione, W.H. Karasov, S. Morzunov, E. Otteson, and S.St. Jeor. (1998) Prevalence of Hanta virus in four species of Neotoma from Arizona and Utah.  Journal of Mammalogy 79:1254-1259.

McWilliams, S.R. and W.H. Karasov (1998) Test of a digestion optimization model: effects of costs of feeding on digestive parameters.  Physiological Zoology 71:168-178

Lepczyk, C. M., Caviedes‑Vidal, E., and Karasov, W. H. (1998)  Digestive responses during food restriction and realimentation in nestling house sparrows (Passer domesticus).  Physiological. Zoology 71:561-573.

McWilliams, S.R., E. Caviedes-Vidal, and W.H. Karasov (1999)  Digestive adjustments in Cedar Waxwings to high feeding rate.  Journal of Experimental Zoology 283:394-407.

Jofre, M.B. and W.H. Karasov (1999).  The direct effect of ammonia on three species of North American anuran amphibians.  Environmental Toxicology and Chemistry 18:1806-1812.

Huang, Y.W., W.H. Karasov, K.P. Patnode, and C.R. Jefcoate (1999). Exposure of northern leopard frogs in the Green Bay ecosystem to polychlorinated biphenyls, polychlorinated dibenzo-p-dioxins, and polychlorinated dibenzofurans is measured by direct chemistry but not hepatic ethoxyresorufin-O-deethylase activity. Environmental Toxicology and Chemistry 18:2123-2130.

Rosenshield, M.L., M.B. Jofre, and W.H. Karasov (1999).  Effects of PCB 126 on green frog (Rana clamitans) and leopard frog (Rana pipiens) hatching success, development, and metamorphosis.  Environmental Toxicology and Chemistry 18:2478-2486.

Lepczyk, C.A. and W.H. Karasov (2000). The effect of ephemeral food restrictions on the growth of house sparrows (Passer domesticus). Auk 117:164-174.

Huang, Y.W. and W.H. Karasov (2000) Oral bioavailability and toxicokinetics of 3,3′,4,4′,5-pentachlorobiphenyl in northern leopard frogs, Rana pipiens.  Environmental Toxicology and Chemistry 19(7):1788-1794.

Karasov, W.H. and M.W. Meyer (2000) Testing the role of contaminants in depressing avian numbers. Revista Chilena de Historia Natural 73:243-252.

Caviedes-Vidal, E., D. Afik, C. Martinez del Rio, and W.H. Karasov (2000) Dietary modulation of intestinal enzymes of the house sparrow (Passer domesticus): testing an adaptive hypothesis.  Comparative Biochemistry and Physiology 125:11-24.

Dearing, M. D., A.M. Mangione, and W.H. Karasov (2000)  Diet breadth of mammalian herbivores: tests of nutrient constraints and detoxification limitations hypotheses. Oecologia 123:397-405.

Mangione, A.M., M.D. Dearing, and W.H. Karasov (2000)  Interpopulation differences in tolerance to creosote bush resin in desert woodrats (Neotoma lepida).  Ecology 81:2067-2076.

Karasov, W.H. and B. Pinshow (2000)  Test for physiological limitation to nutrient assimilation in a long-distance migrant at a springtime stopover site.  Physiological and Biochemical Zoology 73:335-343.

Starck, M., W.H. Karasov, and D. Afik (2000) Intestinal nutrient uptake measurements and tissue damage. Validating the everted sleeves method. Physiological and Biochemical Zoology 73:454-460.

Jofré, M.G., M.L. Rosenshield, and W.H. Karasov (2000) Effects of PCB 126 and ammonia, alone and in combination, on green frog (Rana clamitans) and leopard frog (R. pipiens) hatching success, development, and metamorphosis.  Jour. Iowa Acad. Sci. 107(3):113-122..

Allran, J.W. and W.H. Karasov (2000)  Effects of atrazine and nitrate on northern leopard frog (Rana pipiens) larvae exposed in the laboratory from post-hatch through metamorphosis.  Environmental Toxicology and Chemistry 19(11):2850-2855.

Huang, Y. W., J. J. Stegeman, B. R. Woodin, and W. H. Karasov (2001).  Immunohistochemical localization of cytochrome P450-associated monooxygenases induced by 3,3′,4,4′,5-pentachlorobiphenyl in multiple organs of northern leopard frog, Rana pipiens. Environmental Toxicology and Chemistry 20(1):191-197

Chediack, J.G., E. Caviedes-Vidal, W.H. Karasov, and M. Pestchanker (2001) Passive absorption of hydrophilic carbohydrate probes by house sparrows, Passer domesticus.  Journal of Experimental Biology 204:723-731.

McWilliams, S.R. and W.H. Karasov (2001) Phenotypic flexibility in digestive system structure and function in migratory birds and its ecological significance. Comparative Biochemistry and Physiology A 128:579-593.

Dykstra, C.R., W.H. Karasov, M.W. Meyer, and D.K. Warnke (2001) Daily energy expenditure of nestling bald eagles in northern Wisconsin.  Condor 103:175-179.

Allran, J.W. and W.H. Karasov (2001)  Effects of atrazine on embryos, larvae, and adults of anuran amphibians.  Environmental Toxicology and Chemistry 20:769-775.

Caviedes-Vidal, E. and W.H. Karasov.  (2001) Developmental changes in digestive physiology of nestling house sparrows, Passer domesticus.  Physiological and Biochemical Zoology 74:769-782.

Ciminari, M.E., D. Afik, W.H. Karasov, and E. Caviedes-Vidal (2001)  Is diet-shifting facilitated by modulation of pancreatic enzymes? Test of an adaptational hypothesis in yellow-rumped warblers. Auk 118:1101-1107.

Dearing, M.D., A.M. Mangione, and W.H. Karasov (2001) Plant Secondary Compounds as Diuretics: An Overlooked Consequence of PSC.  American Zoologist 41:890-901.

Mangione, A.M., M.D. Dearing, and W.H. Karasov (2001).  Detoxification processes in relation to toxin tolerance in desert woodrats eating creosote bush. Journal of Chemical Ecology 27:2559-2578.

McWilliams, S. R., S. B. Kearney, W. H. Karasov (2002)  Dietary preferences of warblers for specific fatty acids in relation to nutritional requirements and digestive capabilities.  Journal of Avian Biology 33:167-174.

Dearing, M.D., A.M Mangione, and W.H. Karasov (2002) Ingestion of plant secondary compounds causes diuresis in woodrat herbivores.  Oecologia 130:576-584.

Lee, K.A., W.H. Karasov, and E. Caviedes-Vidal (2002)  Digestive response to restricted feeding in migratory yellow-rumped warblers (Dendroica coronata).  Physiological and Biochemical Zoology 75:314-323.

Fournier, F., W.H. Karasov, K. Kenow, M.W.Meyer, and R.K. Hines (2002). The daily energy expenditures of free-ranging common loon (Gavia immer) chicks.  The Auk 119:1121–1126.

Fournier, F., W.H. Karasov, K.P. Kenow, M.W. Meyer, and R.K. Hines (2002).  The oral bioavailability and toxicokinetics of methylmercury in common loon (Gavia immer) chicks.  Comparative Biochemistry and Physiology A 133:703-714.

Warnke, D.K., D.E. Andersen, C.R. Dykstra, M.W. Meyer, and W.H. Karasov.  (2002) Provisioning rates and time budgets of adult and nestling Bald Eagles at inland Wisconsin nests.  Journal of Raptor Research 36:121-127.

Kenow, K.P., S. Gutreuter, R.K. Hines, M.W. Meyer, F. Fournier, and W.H. Karasov (2003).  Effects of methyl mercury exposure on the growth of juvenile Common loons.  Ecotoxicology 12:171-182.

Kenow, K.P., M.W. Meyer, F. Fournier, W.H. Karasov, A. Elfessi, and S. Gutreuter (2003) Effects of subcutaneous transmitter implants on behavior, growth, energetics, and survival of Common loon chicks. Journal of Field Ornithology 74:179-186.

Chediack, J.G., E. Caviedes-Vidal, V. Fasulo, J. Yamin, and W.H. Karasov (2003).  Intestinal passive absorption of hydrosoluble compounds by sparrows: effects of molecular size and luminal nutrients.  Journal of Comparative Physiology B 173:187-197.

Chang, M.H., J.G. Chediack, E. Caviedes-Vidal, and W.H. Karasov.  (2004)  L-glucose absorption in house sparrows (Passer domesticus) is nonmediated. Journal of Comparative Physiology B 174:181-188.

Karasov, W.H., B. Pinshow, J.M. Starck, D. Afik (2004) Anatomical and histological changes in the alimentary tract of migrating blackcaps (Sylvia atricapilla): a comparison among fed, fasted, food-restricted and refed birds.  Physiological and Biochemical Zoology 77:149-160.

Chang, M. H. and W. H. Karasov. (2004). Absorption and paracellular visualization of fluorescein, a hydrosoluble probe, in intact house sparrows (Passer domesticus). Zoology 107:121-133.

Mangione, A.M., M.D. Dearing, and W.H. Karasov. (2004)  Creosote bush (Larrea tridentata) resin increases water demands and reduces energy availability in desert woodrats (Neotoma lepida).  Journal of Chemical Ecology 30:1409-1429.

Chang, M.H. and W.H. Karasov. (2004) How the house sparrow Passer domesticus absorbs glucose (2004) Journal of Experimental Biology 207:3109-3121.

Skopec, M., A. Hagerman, and W.H. Karasov. (2004) Do salivary proline-rich proteins counteract dietary hydrolysable tannin in laboratory rats?  Journal of Chemical Ecology 30:1679-1692.

Jung, R.E., W.H. Karasov, M. J. Melancon.  (2004)  Cytochrome P450 activity in green frogs (Rana clamitans melanota) exposed to water and sediments in the Fox River and Green Bay, Wisconsin.  Bulletin of Environmental Contamination and Toxicology 73(6): 955-962.

Green, A.K., S. Haley, M.D. Dearing, D.M. Barnes, and W.H. Karasov (2004)  Intestinal capacity of P-glycoprotein is higher in the juniper specialist, Neotoma stephensi, than the sympatric generalist, N. albigula.  Comparative Biochemistry and Physiology A 139:325-333.

Green, A.K., D.M. Barnes, and W.H. Karasov  (2005)  A new method to measure intestinal activity of P-glycoprotein in avian and mammalian species.  Journal of Comparative Physiology B: 175 (1): 57 – 66.

Karasov, W.H., R.E. Jung, S. VandenLangenberg, and T.E. Bergeson (2005)  Field exposure of frog embryos and tadpoles along a pollution gradient in the Fox River and Green Bay ecosystem in Wisconsin.  Environmental Toxicology and Chemistry 24(4): 942-953.

McWhorter, T.J., B.H. Bakken, W.H. Karasov, and C. Martinez del Rio (2006) Hummingbirds rely on both paracellular and carrier-mediated intestinal glucose absorption to fuel high metabolism.  Biology Letters 2:131-134.

Chediack, J.G., E. Caviedes-Vidal, and W.H. Karasov (2006) Electroaffinity in paracellular absorption of hydrophilic D-dipeptides by sparrow intestine.  Journal of Comparative Physiology 176(4):303-309

Chen, T.H., J.A. Gross, and W.H. Karasov (2006) Sublethal effects of lead on northern leopard frog (Rana pipiens) tadpoles. Environmental Toxicology and Chemistry 25(5):1383-1389.

Fassbinder-Orth, C. and W.H. Karasov (2006) Effects of feed restriction and realimentation on digestive and immune function in the leghorn chick. Poultry Science 85:1449-1456.

Green, A.K., S.L. Haley, D.M. Barnes, M.D. Dearing, and W.H. Karasov (2006) Is alpha-pinene a substrate for permeability-glycoprotein in woodrats? Journal of Chemical Ecology 32(6):1197-1211.

Huang, Y.W., D.J. Hoffman, and W.H. Karasov (2007)  Oxidative stress induced in PCB 126-exposed Northern leopard frogs, Rana pipiens. Journal of Toxicology and Environmental Health Part A 70:676-681.

Karasov, W.H., K.P. Kenow, M.W. Meyer, and F. Fournier (2007) Bioenergetic and pharmacokinetic model for exposure of Common Loon chicks to methylmercury.  Environmental Toxicology and Chemistry 26: 67-685.

Tracy, C.R., T.J. McWhorter, C. Korine, M. Wojciechowski, B. Pinshow, and W.H. Karasov (2007) Absorption of sugars in the Egyptian fruit bat (Rousettus aegyptiacus): a paradox explained.  Journal of Experimental Biology 210:1726-1734.

McWhorter, T.J. and W.H. Karasov (2007) Paracellular nutrient absorption in a gum-feeding New World primate, the common marmoset Callithrix jacchus.  American Journal of Primatology 69:1-13.

Kenow, K.P., M.W. Meyer, R.K. Hines, and W.H. Karasov (2007)  Distribution and accumulation of mercury in tissues of captive-reared common loon (Gavia immer) chicks.  Environmental Toxicology and Chemistry 26:1047-1055.

Lavin, S.R., T.J. McWhorter, and W.H. Karasov (2007) Mechanistic bases for differences in passive absorption. Journal of Experimental Biology 210:2754-2764.

Fournier, F. W.H. Karasov, K.P. Kenow, and M.W. Meyer (2007)  Growth and energy requirements of captive-reared common loon chicks (Gavia immer).  The Auk 124(4):1158-1167.

Gross, J.A., T.H. Chen, and W.H. Karasov (2007) Lethal and sublethal effects of chronic cadmium exposure on northern leopard frog (Rana pipiens) tadpoles. Environmental Toxicology and Chemistry 26:1192-1197.

Chen, T.H., J.A. Gross, and W.H. Karasov (2007) Adverse effects of chronic copper exposure in larval northern leopard frogs (Rana pipiens).  Environmental Toxicology and Chemistry 26:1470-1475.

Caviedes-Vidal, E., T.J. McWhorter, S.R. Lavin, J.G. Chediack, C.R. Tracy, and W.H. Karasov (2007) The digestive adaptation of flying vertebrates: high intestinal paracellular absorption compensates for smaller guts.  Proceedings of the National Academy of Science, USA 104(4):19132-19137.

Naya, D.E., W.H. Karasov, and F. Bozinovic (2007) Phenotypic plasticity in laboratory mice and rats: a meta-analysis of current ideas on gut size flexibility.  Evolutionary Ecology Research 9:1363-1374.

Caviedes-Vidal, E., W.H. Karasov, J.G. Chediack, V. Fasulo, P.A. Cruz-Neto, and L. Otani. (2008) Paracellular absorption: a bat breaks the mammal paradigm. PLoS ONE 3(1): e1425. doi:10.1371/journal.pone.0001425.

Jofre, M.G. and W.H. Karasov (2008) Effect of mono-ortho and di-ortho substituted polychlorinated biphenyl (PCB) congeners on leopard frog survival and sexual development. Chemosphere 71:1609-1619.

Tsurim, I., N. Sapir, J. Belmaker, I. Shanni, I. Izhaki, M.S. Wojciechowski, W.H. Karasov, and B. Pinshow (2008) Drinking water boosts food intake rate, body mass increase and fat accumulation in migratory blackcaps (Sylvia atricapilla).  Oecologia 156:21–30.

Lavin, S.R. and W.H. Karasov (2008) Allometry of paracellular absorption in birds.  Physiological and Biochemical Zoology 81(5):551-560.

Lavin, S.R., W.H. Karasov, A.R. Ives, K.M. Middleton, and T. Garland, Jr. (2008) Morphometrics of the avian small intestine, compared with non-flying mammals: A phylogenetic approach. Physiological and Biochemical Zoology 81(5):526-550.

Naya, D.E., W.H. Karasov, and F. Bozinovic (2008) Latitudinal trends in digestive flexibility: testing the climatic variability hypothesis with data on the intestinal length of rodents.  American Naturalist 172(4):E122-E134.

Gross, J.A., P.T.J. Johnson, L.K. Prahl, and W.H. Karasov (2009) Critical period of sensitivity for effects of cadmium on frog growth and development. Environmental Toxicology and Chemistry 28(6):1227-1232.

Gilbert, J.H., P.A. Zollner, A.K. Green, J.L. Wright, and W.H. Karasov (2009).  Seasonal field metabolic rates of American martens in Wisconsin. American Midland Naturalist 169:330-337.

Brzęk, P., K. Kohl, E. Caviedes-Vidal, and W.H. Karasov (2009) Developmental adjustments of House sparrow (Passer domesticus) nestlings to diet composition. Journal of Experimental Biology 212:1284-1293.

Karasov,W.H. and H.V. Carey (2009) Metabolic teamwork between gut microbes and hosts. Microbe 4(7):323-328.

McWhorter,T.J., E. Caviedes-Vidal, and W.H. Karasov (2009) The integration of digestion and osmoregulation in the avian gut.  Biological Reviews 84(4):553-565 ( doi:10.1111/j.1469-185X.2009.00086.x)

Chen, T-H, J.A. Gross, and W.H. Karasov (2009) Chronic exposure to pentavalent arsenic of larval leopard frogs (Rana pipiens): Bioaccumulation and reduced swimming performance.  Ecotoxciology (DOI10.1007/s10646-009-0316-3).

Fassbinder-Orth, C.A., E.K. Hofmeister, C.Weeks-Levy and W.H. Karasov (2009)  Oral and parenteral immunization of chickens (Gallus gallus) against West Nile virus with recombinant envelope protein. Avian Diseases: Vol. 53, No. 4, pp. 502–509 (doi: 10.1637/8688-031009-Reg.1)

McWhorter, T.J., A.K. Green, and W.H. Karasov (2010) Assessment of radiolabeled D-glucose and the non-metabolizable analog 3-O-methyl-D-glucose as tools for in vivo absorption studies.  Physiological and Biochemical Zoology 83(2): 376-384. DOI:10.1086/597524

Cary Coyle, T.L. and W.H. Karasov (2010) Chronic dietary polybrominated diphenyl ether (PBDE) exposure affects survival, growth and development of Rana pipiens tadpoles. Environmental Toxicology and Chemistry 29(1):133-141.

Tracy, C.R., T.J. McWhorter, M.S. Wojciechowski, B. Pinshow, and W.H. Karasov (2010)  Carbohydrate absorption by blackcap warblers (Sylvia atricapilla) changes during migratory refueling stopovers. Journal of Experimental Biology 213(3): 380-385.

Brzęk, P., K.M. Lessner, E. Caviedes-Vidal, and W.H. Karasov (2010) Low plasticity in digestive physiology constrains feeding ecology in diet specialist, Zebra finch (Taeniopygia guttata). Journal of Experimental Biology 213(5):798-807.

Skopec, M.M., A.K. Green, and W.H. Karasov (2010) Flavonoids have differential effects on glucose absorption in rats (Rattus norvegicus) and American robins (Turdus migratorius).  Journal of Chemical Ecology 36:236-243.

Brzęk,P., E.Caviedes-Vidal, K. Hoefer, and W.H. Karasov (2010) Effect of age and diet on total and passive glucose absorption in nestling House sparrows.  Physiological and Biochemical Zoology 83(3):501-511.

Wright, J., W.H. Karasov, A.M.N. Kazem, I.B. Goncalves, and E. McSwan (2010) Begging and digestive responses to differences in long-term and short-term need in nestling pied flycatchers. Animal Behaviour 80:517-525.

Liu, J.N. and W.H. Karasov (2011) Hibernation in warm hibernacula by free ranging Formosan Leaf-Nosed Bats, Hipposideros terasensis, in subtropical Taiwan. Journal of Comparative Physiology, B. Biochemical, Systematic, and Environmental 181:125-135 (DOI10.1007/s00360-010-0509-3).

Karasov, W.H., C. Martinez del Rio, and E. Caviedes-Vidal (2011) Ecological physiology of diet and digestive systems. Annual Review of Physiology 73:69–93.

Kohl, K.D., P. Brzęk, E. Caviedes-Vidal, and W.H. Karasov (2011) Pancreatic and intestinal carbohydrases are matched to dietary starch level in wild passerine birds.  Physiological and Biochemical Zoology 84:195-203.

Naya, D.E., and W.H. Karasov (2011) Food digestibility by microbes in wild ruminants: the effect of host species and dietary substrate. Rangelands 33(1):31-34.

Brzek, P., K.D. Kohl, E. Caviedes-Vidal, and W.H. Karasov (2011)  Fully reversible phenotypic plasticity of digestive physiology in young House sparrows: lack of long-term effect of diet composition. Journal of Experimental Biology 214:2755-2760.

Karasov, W.H. (2011)  Digestive physiology: a view from molecules to ecosystem.  American Journal of Physiology 301:R276-R284.

Olivia E. LeDee, O.E., W.H. Karasov, K.J. Martin, M.W. Meyer, C.A. Ribic, and T.R. Van Deelen (2011)  Envisioning the future of wildlife in a changing climate: collaborative learning for adaptation planning. Wildlife Society Bulletin 35(4):508-513; DOI: 10.1002/wsb.62

Liu, J.N and W.H. Karasov (2012) Metabolism during winter in a subtropical hibernating bat, the Formosan leaf-nosed bat (Hipposideros terasensis). Journal of Mammalogy 93:220-228; DOI: 10.1644/11-MAMM-A-144.1.

Van Schmidt, N.D., T.L. Cary, M.E. Ortiz-Santaliestra, and W.H. Karasov (2012) Effects of chronic polybrominated diphenyl ether (PBDE) exposure on gondal development in the northern leopard frog, Rana pipiens. Environmental Toxicology and Chemistry 31:347-354. DOI: 10.1002/etc.1702

Karasov, W.H., E. Caviedes-Vidal, B. Hartman Bakken, I. Izhaki, M. Samuni-Blank, and Z. Arad. (2012) Capacity for absorption of water-soluble secondary metabolites greater in birds than in rodents.  PLoS ONE 72(1) e32417. doi:10.1371/journal.pone.0032417.

Killpack, T.L. and W.H. Karasov (2012) Growth and development of house sparrows (Passer domesticus) in response to chronic food shortage throughout the nestling period.  Journal of Experimental Biology 215:1806-1815.

Samuni-Blank, M., I. Izhaki, M.D. Dearing, Y. Gerchman, B. Trablshy, A. Lotan, W.H. Karasov, Z. Arad (2012) Intraspecific directed deterrence by the mustard oil bomb in a desert plant. Current Biology 22:1-3.

Killpack, T.L. and W.H. Karasov (2012) Ontogeny of adaptive antibody response to a model antigen in captive altricial Zebra finches. PLoS ONE 7(10) e47294.

Fasulo, V., Z.Q. Zhang, J.G. Chediack, F.D. Cid, W.H. Karasov and E. Cavides-Vidal (2013) The capacity for paracellular absorption in the insectivorous bat Tadarida brasiliensis. Journal of Comparative Physiology B 183(2):289-296. DOI 10.1007/s00360-012-0696-1.

Fasulo, Veronica, Z.Q. Zhang, E.R., J.G. Chediack, W.H. Karasov, and E. Caviedes-Vidal (2013).  Paracellular absorption in laboratory mice: molecule size-dependent but low capacity. Comparative Biochemistry and Physiology, Part A 164:71–76.

Price, E.R., A. Brun, V. Fasulo, W.H. Karasov, and E. Caviedes-Vidal (2013)  Intestinal perfusion indicates high reliance on paracellular nutrient absorption in an insectivorous bat Tadarida brasiliensis. Comparative Biochemistry and Physiology, Part A 164: 351–355 (http://dx.doi.org/10.1016/j.cbpa.2012.11.005).

McWhorter, T.J., B. Pinshow, W.H. Karasov, and C.R. Tracy (2013). Paracellular absorption is relatively low in the herbivorous Egyptian Spiny-tailed Lizard, Uromastix aegyptia. PLoS ONE 8(4): e61869. doi:10.1371/journal.pone.0061869.

Brzęk, P., E. Ciminari, K.D. Kohl, K. Lessner, E. Caviedes-Vidal, and W.H. Karasov (2013) Effect of age and diet composition on activity of pancreatic enzymes in birds. Journal of Comparative Physiology B 183:685-697 DOI 10.1007/s00360-012-0731-2.

Karasov, W.H. and A.E. Douglas (2013) Comparative Digestive Physiology. Comprehensive Physiology 3:741-783.

Samuni-Blank, M., Z. Arad, M.D. Dearing, Y. Gerchman, W.H. Karasov, and I. Izhaki. (2013) Friend or foe? Disparate plant-animal interactions in two congeneric rodents. Evolutionary Ecology DOI 10.1007/s10682-013-9655-x.

Meier, S.A., Fassbinder-Orth, C.A., and W.H. Karasov (2013).  Ontogenetic changes in innate immune function in captive and wild subspecies of prairie chickens (Tympanuchus cupido spp.). Journal of Wildlife Management 77(3):633-638.

Cary, T.L. and W.H. Karasov (2013) Toxicokinetics of polybrominated diphenyl ethers across life stages in the northern leopard frog, Lithobates pipiens.  Environmental Toxicology and Chemistry 32(7):1631-1640.

Ortiz-Santaliestra, M.E., T.A.G. Rittenhouse, T.L. Cary, and W.H. Karasov (2013).  Interspecific and post-metamorphic variation in susceptibility of three North American anurans to Batrachochytrium dendrobatidis.  Journal of Herpetology 47(2):286-292.

Killpack, T.L., Y. Oguchi, and W.H. Karasov (2013) Ontogenetic patterns of constituitive innate immune parameters in altricial house sparrow nestlings. Journal of Avian Biology 44:513-520 [Editor’s Choice].

Samuni-Blank, M., I. Izhaki, M.D. Dearing, W.H. Karasov, Y. Gercham, K.D. Kohl, P. Lymberakis, P. Kurnath, and Z. Arad (2013) Physiological and behavioral effects of fruit toxins on seed-predating versus seed-dispersing congeneric rodents. Journal of Experimental Biology 216:3667-3673.

Brzek, P., K.D. Kohl, E. Caviedes-Vidal, and W.H. Karasov (2013) Activity of intestinal carbohydrases responds to multiple dietary signals in nestling House sparrows. Journal of Experimental Biology 216:3981-3987.

Price, E.R., L.J. Ruff, A. Guerra, and W.H. Karasov (2013) Cold exposure increases paracellular permeability to nutrients in the mouse. Journal Experimental Biology 216:4065-4070.

Kohl, K.D., T.L. Cary, W.H. Karasov, and M.D. Dearing (2013) Restructuring of the amphibian gut microbiota through metamorphosis. Environmental Microbiology and Environmental Microbiology Reports 5(6) 899-903.

McWilliams, S.R. and W.H. Karasov (2014) Spare capacity and phenotypic flexibility in the digestive system of a migratory bird: defining the limits of animal design. Proceedings of the Royal Society B 281: 20140308. http://dx.doi.org/10.1098/rspb.2014.0308.

Killpack, T.L., D.N. Tie, and W.H. Karasov (2014)  Compensatory growth in nestling Zebra Finches impacts body composition but not adaptive immune function.  The Auk 131:396-406.

Cary, T.L., M.E. Ortiz-Santaliestra, and W.H. Karasov. (2014) Immunomodulation in post-metamorphic northern leopard frogs, Lithobates pipiens, following larval exposure to polybrominated diphenyl ether. Environmental Science & Technology 48(10): 5910-5919.  dx.doi.org/10.1021/es405776m

Brun, A., E.R. Price, M.N. Gonter-Fourcade, G. Fernandez-Marinone, A.P. Cruz-Neto, W.H. Karasov, and E. Caviedes-Vidal (2014) High paracellular nutrient absorption in intact bats is associated with high paracellular permeability in perfused intestinal segments. Journal of Experimental Biology 217:3311-3317.  doi:10.1242/jeb.104927

Price, E.R., K.H. Rott, E. Caviedes-Vidal, and W.H. Karasov (2014) Paracellular nutrient absorption is higher in bats than rodents: integrating from intact animals to the molecular level. Journal of Experimental Biology 217:1-10. doi:10.1242/jeb.105619

Samuni-Blank, M., I. Izhaki, Y. Gerscham, M.D. Dearing, W.H. Karasov, B. Travelc, T. Edwards, Z. Arad (2014) Taste and physiological responses to glucosinolates: Seed predator versus seed disperser. PLoS ONE 9(11):e112505.

Ferrie, G.M., V. C. Alford, J. Atkinson, E. Baitchman, D. Barber, W. S. Blaner, G. Crawshaw, A. Daneault, E. Dierenfeld, M. Finke, G. Fleming, R. Gagliardo, E. A. Hoffman, W.H. Karasov, K. Klasing, E. Koutsos, J. Lankton, S.R. Lavin, A. Lentini, S. Livingston, B. Lock, T. Mason, A. McComb, C. Morris, A. P. Pessier, F. Olea‐Popelka, T. Probst, C. Rodriguez, K. Schad, K. Semmen, J. Sincage, M. A. Stamper, J. Steinmetz, K. Sullivan, S. Terrell, N. Wertan, C. J. Wheaton, B. Wilson, and E. V. Valdes (2014) Nutrition and health in amphibian husbandry. Zoo Biology 33(6):485-501

Zhang, Z, A. Brun, E. Price, A.P. Cruz-Neto, W.H. Karasov, E. Caviedes-Vidal (2015) A comparison of mucosal surface area and villous histology in small intestines of the Brazilian free-tailed bat (Tadarida brasiliensis) and the mouse (Mus musculus). Journal of Morphology 276:102-108. DOI 10.1002/jmor.20324.

Richman,S.E., J.O. Leafloor, W.H. Karasov, and S.R. McWilliams (2015) Ecological implications of reduced forage quality on growth and survival of sympatric geese. Journal of Animal Ecology 84(1): 284-298. doi: 10.1111/1365-2656.12270.

Price, E.R., A. Brun, E. Caviedes-Vidal, and W.H. Karasov (2015) Digestive adaptations of aerial lifestyles. Physiology 30:69-78.   doi:10.1152/physiol.00020.2014

Kohl, K.D., T.L. Cary, W.H. Karasov, and M.D. Dearing (2015)  Larval exposure to polychlorinated biphenyl 126 (PCB-126) causes persistent alteration of the amphibian gut microbiota. Environmental Toxicology and Chemistry 34(5):1113-1118.

Killpack, T.L., E. Carrel, and W.H. Karasov (2015) Impacts of short-term food restriction on immune development in altricial house sparrow nestlings. Physiological and Biochemical Zoology 88(2):195-207.

Price, E.R., A. Brun, M. Gontero-Fourcade, G. Fernandez-Marinone, A.P. Cruz-Neto, W.H. Karasov, and E. Caviedes-Vidal. (2015) Intestinal water absorption varied with expected dietary water load among bats but does not drive paracellular nutrient absorption.  Physiological and Biochemical Zoology 88(6):680-684.

Lessner, K.M., M. Denise Dearing, I. Izhaki, M. Samui-Blanck, Z. Arad, and W.H. Karasov (2015) Small intestinal hydrolysis of plant glucosides: higher glucohydrolase activities in rodents than passerine birds. Journal of Experimental Biology 218, 2666-2669. doi:10.1242/jeb.121970

Gatica-Sosa,C., Brzek,P., Chediack,J.G., Cid, F.D., Karasov,W.H., Caviedes-Vidal, E. (2016) Differential transcriptional responses underlie dietary induction of intestinal carbohydrase activities in house sparrow nestlings. Journal of Animal Physiology and Animal Nutrition 100: (2016) 236-242.  DOI: 10.1111/jpn.12354.

Price, E.R., K.H. Rott, E. Caviedes-Vidal, and W.H. Karasov (2016) Claudin gene expression patterns do not associate with interspecific differences in paracellular nutrient absorption. Comparative Biochemistry and Physiology Part B: Biochemistry and Molecular Biology 191:36-45.

Kohl,K.D., M. Samuni-Blank, P. Lymberakis, P. Kurnath, I. Izhaki, Z. Arad, W.H. Karasov, and M.D. Dearing (2016) Effects of fruit toxins on intestinal and microbial bglucosidase activities of seed-predating and seed-dispersing rodents. Physiological and Biochemical Zoology 89(3):198-205.

Rodriguez-Peña, N., E.R. Price, E. Caviedes-Vidal, C.M. Flores-Ortiz, and W.H. Karasov (2016) Intestinal paracellular absorption is necessary to support the sugar oxidation cascade in nectarivorous bats. Journal of Experimental Biology 219:779-782. Editor’s ChoiceVol. 219, No. 6.

Gleason, S.M., J.M. Yahn, and W.H. Karasov (2016) Digestive efficiency of Northern Leopard Frog (Lithobates pipiens) during development, reared on a laboratory diet.  Herpetologica 72(2):107-113. doi: 10.1655/HERPETOLOGICA-D-15-00028

DeQuattro, Z.A., and W.H. Karasov (2016) Impacts of 2,4-Dichlorophenoxyacetic acid aquatic herbicide formulations on reproduction and development of the fathead minnow (Pimephales promelas). Environmental Toxicology and Chemistry 35(6):1478-1488.

Pauli, J.N., M.Z. Peery, E.D. Fountain, and W.H. Karasov (2016) Arboreal folivores limit their energetic output, all the way to slothfulness. American Naturalist 188(2): 196-204 DOI: 10.1086/687032

Freitas, M.B., C.A. Tsai, and W.H.Karasov (2017) Warmer temperature modifies the effect of polybrominated biphenyl ethers (PBDEs) on hormone profiles during metamorphosis in Lithobates pipiens.  Environmental Toxicology and Chemistry 36 (1) 120-127.

Kohl, K.D., M.E. Ciminari, J.G. Chediack, J.O. Leafloor, W.H. Karasov, S.R. McWilliams, and E. Caviedes-Vidal (2017)  Modulation of digestive enzyme activities in the avian digestive tract in relation to diet composition and quality. Journal of Comparative Physiology 187(2):339-351.

Cornelius, E.A., F. Vezina, L. Regimbald, F. Hallot, M. Petit, O. Love, and W.H. Karasov (2017) Chickadees faced with unpredictable food increase fat reserves but certain components of their immune function decline. Physiological and Biochemical Zoology 90(2): 190-200.  DOI:10.1086/689913.

Rott, K.H., E. Caviedes-Vidal, and W.H. Karasov (2017) Intestinal digestive enzyme modulation in house sparrow nestlings occurs within 24 hours of a change in diet composition.  Journal of Experimental Biology 220:2733-2742.

Karasov, W.H. (2017) Integrative physiology of trans- and paracellular intestinal absorption.  Journal of Experimental Biology 220:2495-2501.

Fathi, F., A. Brun, K.H. Rott, P.F. Cobra, M. Tonelli, H.R. Eghbalnia, E. Caviedes-Vidal, W.H. Karasov and J.L. Markley (2017) NMR-based identification of metabolites in polar and non-polar extracts of avian liver. Metabolites 2017, 7, 61; doi:10.3390/metabo7040061.

Kohl, K.D., A. Brun, K.H. Rott, S.R. Bordenstein, E. Caviedes-Vidal, and W.H. Karasov (2018)  Gut microbes limit growth in House sparrow nestlings (Passer domesticus), but not through limitations in digestive capacity. Integrative Zoology 13:139-151.

Gatica-Sosa, C. P. Brzek, M. Magallanes, W.H. Karasov, and E. Caviedes-Vidal (2018) Intestinal α –glucosidase transcriptional responses during development and diet adjustment in altricial birds. Journal of Experimental Biology 221(6):jeb171827.

Latimer, C., S. Cooper, W.H. Karasov, and B. Zuckerberg (2018) Does habitat fragmentation promote climate-resilient phenotypes?  Oikos 127(8)1069-1080  (Editor’s Choice, Aug. 2018 Issue).

Garro, C., A. Brun, W.H. Karasov, and E. Caviedes-Vidal (2018) Small intestinal epithelial permeability to water soluble nutrients higher in passerine birds than in rodents.  Journal of Animal Physiology and Animal Nutrition 102(6):1766-1773.

Dehnert, G.K., M.F. Freitas, Z.A. DeQuattro, T. Barry and W.H. Karasov (2018) Effects of low, subchronic exposure of 2,4-Dichlorophenoxyacetic acid (2,4-D) and commercial 2,4-D formulations on early life stages fathead minnows (Pimephales promelas). Environmental Toxicology and Chemistry 37(10):2550-2559.

Kohl, K.D., A. Brun, E. Caviedes-Vidal, and W.H. Karasov (2019) Age-related changes in the gut microbiota of  wild House sparrow nestlings. Ibis 161:184-191 (doi 10.1111/ibi.12618).

Bourne, A.R., A.E. McKechnie, S.J. Cunningham, A.R. Ridley, S.M. Woodborne, and W.H. Karasov (2019) Non-invasive measurement of metabolic rates in wild, free-living birds using doubly labelled water. Functional Ecology 33(1) :162-175 (DOI: 10.1111/1365-2435.13230).

Cornelius-Ruhs, E., F. Vezina, and W.H. Karasov (2019) Physiological and immune responses of free-living, temperate birds provided a gradient of food supplementation. Physiological and Biochemical Zoology 92(1):106-114.

Dehnert, G.K., W.H. Karasov, and M.A. Wolman (2019) 2,4-Dichlorophenoxyacetic acid containing herbicide impairs essential visually guided behaviors of larval fish. Aquatic Toxicology 209 (2019): 1-12. Share Link to preview: https://authors.elsevier.com/c/1YRw6,3oDQcUjT

Wang, L., E. Caviedes-Vidal, and W.H. Karasov (2019) Diet composition modulates intestinal hydrolytic enzymes in white-footed mice (Peromsycus leucopus). Journal of Mammalogy DOI:10.1093/jmammal/gyz110.

Brun, A., G.F. Marinone, E.R. Price, L.A. Nell, B. Marqus, A. Castellar, M. Gontero-Fourcade, A. P. Cruz-Neto, W.H. Karasov, and E. Caviedes-Vidal (2019).  Morphological bases for intestinal paracellular absorption in bats and rodents. Journal of Morphology DOI:10.1002/jmor.21037.

Cornelius-Ruhs, W., F. Vézina, M.A. Walker, and W.H. Karasov (2019) Who pays the bill? The effects of altered brood size on parental and nestling physiology. Journal of Ornithology (in press). DOI 10.1007/s10336-019-01715-1

Brun, A., M.E. Magallanes, C. Martinez del Rio, G.A. Barrett-Wilt, W.H. Karasov, and E. Caviedes-Vidal (2020). A fast and accurate method to identify and quantify enzymes in brush-border membranes: In situ hydrolysis followed by nano LC-MS/MS. Methods and Protocols 3, 15; doi:10.3390/mps3010015.

Brun, A., D. Mendez-Aranda, M.E. Magallanes, W.H. Karasov, C. Martinez del Rio, M.W. Baldwin, and E. Caviedes-Vidal (2020) Duplications and functional convergence of intestinal carbohydrate-digesting enzymes. Molecular Biology & Evolution (in press). doi:10.1093/molbev/msaa034

Noakes, J.J., W.H. Karasov, and A. McKechnie (2020) Seasonal variation in body composition in an Afrotropical passerine bird: increases in pectoral muscle mass are, unexpectedly, associated with lower thermogenic capacity. Journal of Comparative Physiology-B (in press).

+Papers submitted to refereed journals, not yet accepted - 1

Yahn, J.M. and W.H. Karasov.  The effects of rearing temperature and dietary PBDE exposure on tadpole growth, development, and their underlying processes. Environmental Toxicology and Chemistry.

+Papers in, or accepted by, non-refereed journals - 3

Diamond, J. M., and W. H. Karasov (1983) Trophic control of the intestinal mucosa.  Nature 304:18.

Karasov, W.H. (2005) Physiological mechanisms of resource use by migrating birds and their conservation implications. Passenger Pigeon 67(1):5-14.

Martinez del Rio, C. & Karasov, W. H. (2010) Body Size and Temperature: Why They Matter. Nature Education Knowledge 1(10):10 (http://www.nature.com/scitable/knowledge/library/body-size-and-temperature-why-they-matter-15157011)

+Invited papers published in conference proceedings - 2

Karasov, W. H. (1991) Ecological physiology of food exploitation:  Insights from measurements and models of digestive function.  In:  Proceedings of the 20th International Ornithological Congress Vol. IV:2159-2169.

Karasov, W. H. (1994) Digestive adaptations in avian omnivores.  Pp. 494-497 in Proceedings of the XV International Congress of Nutrition, Smith-Gordon.

+Book - 1

Karasov, W.H. and C. Martinez del Rio (2007).  Physiological Ecology: How Animals Process Energy, Nutrients, and Toxins. Princeton University Press, Princeton.

+Chapters in Books - 17

Karasov, W. H., R. K. Buddington, and J. M. Diamond (1985) Adaptation of intestinal sugar and amino acid transport in vertebrate evolution.  In Transport processes, iono‑, and osmoregulation, edited by R. Gilles and M. G. Gilles‑Baillien, pp. 227‑239, Springer Verlag, Berlin.

Karasov, W. H., and J. M. Diamond (1987) Adaptation of Intestinal Nutrient Transport.  In Physiology of the Gastrointestinal Tract, edited by L. R. Johnson, Vol. II, pp. 1489-1497, Raven Press, New York.

Karasov, W. H. (1987) Nutrient requirements and the design and function of guts in fish, reptiles, and mammals.  Pp. 181-191 in Comparative Physiology:  Life in Water and on Land, edited by P. Dejours, L. Bolis, C. R. Taylor, E. R. Weibel.  Fidia Research Series, IX-Liviana Press, Padova.

Karasov, W. H. (1988) The nutritional significance of regulation of intestinal nutrient transport.  Pp. 51-63 in Nutritional Modulation of Neural Function, edited by John E. Morley, John H. Walsh and M. Barry Sterman.  Academic Press, New York.

Karasov, W. H. (1988) Nutrient transport across vertebrate intestine.  Pp. 131-172 in Advances in Environmental and Comparative Physiology, Vol. 2, edited by R. Gilles.  Springer-Verlag.

Karasov, W. H. (1989) Adaptation of nutrient transport during vertebrate evolution.  Pp. 157-168 in Adaptation and Development of Gastro-intestinal Function, edited by M. W. Smith and F. V. Sepulveda.  Manchester University Press, Manchester.

Karasov, W. H. (1990) Digestion in birds:  Chemical and physiological determinants and ecological implications.  Pp. 391-415 in M. L. Morrison, C. J. Ralph, J. Verner, and J. R. Jehl, eds.  Avian Foraging:  Theory, Methodology, and Applications.  Studies in Avian Biology No. 13, Cooper Ornithological Society, Kansas.

Meyer, M. W., and W. H. Karasov (1991) Chemical aspects of herbivory in arid and semi-arid habitats.  In:  Plant Chemical Defenses Against Mammalian Herbivory, ed. Palo, R. T. and C. T. Robbins, CRC Press, Inc., Boca Raton, Florida.

Martinez del Rio, C., S. J. Cork, and W. H. Karasov (1994) Modeling gut function:  An introduction.  Pp. 25-53 In:  The Digestive System in Mammals:  Food, Form and Function, edited by D. J. Chivers and P. Langer, Cambridge University Press.

Karasov, W. H. (1996) Digestive plasticity in avian energetics and feeding ecology.  Pp. 61-84 In:  Avian Energetics and Nutritional Ecology, edited by C. Carey, Chapman & Hall, Inc., New York.

Jakubas, W.J., C.G. Guglielmo, and W.H. Karasov (1997). Dilution and detoxication costs: relevance to avian herbivore food selection.  Pp. 53-70 in Mason, J.R., ed. Repellants in Wildlife Management.  Colorado State University Press, Fort Collins, CO.

Vispo, C., and W. H. Karasov (1997) The interaction of avian gut microbes and their host:  An elusive symbiosis.  Ch. 5 in Gastrointestinal Microbiology, edited by R. I. Mackie and B.A. White, pp. 116-155, Chapman & Hall, New York.

Karasov, W. H., and I. D. Hume (1997) The vertebrate gastrointestinal system.  Pp. 407-480 in Handbook of Physiology, Section 13: Comparative Physiology, Vol. 1, edited by W. H. Dantzler, Oxford University Press, New York

Karasov, W.H. and J. Wright (2002)  Nestling digestive physiology and begging.  Pp. 199-219 in Wright, J. and M. Leonard, The Evolution of Begging: Competition, Cooperation, and Communication.  Kluwer Academic Publishers, Dordrecht, The Netherlands.

Karasov, W. H. and S. R. McWilliams. (2005) Digestive constraints in mammalian and avian ecology. Chapter 4, pp 87 – 112 in J. M. Starck and T. Wang, eds. Physiological and Ecological Adaptations to Feeding In Vertebrates, Science Publishers, Inc., Enfield, New Hampshire.

McWilliams, S. R. and W. H. Karasov. (2005) Migration takes guts: digestive physiology of migratory birds and its ecological significance.  Ch. 6, pp 67-78 in P. Mara and R. Greenberg, eds. Birds of Two Worlds. Smithsonian Institution Press, Washington, D.C.

Karasov, W.H. (2012)  Terrestrial vertebrates. Ch. 18 in Metabolic Ecology, R.M. Sibly, J.H. Brown, and A. Kodric-Brown, Wiley-Blackwell.

+Contributed papers and/or abstracts - 137

*Karasov, W. H. (1979) Winter feeding by antelope ground squirrels (Ammospermophilus leucurus):  A case of time limitation.  Bulletin of the Ecological Society of America 60:90.

Anderson, R. A., and W. H. Karasov (1980) Contrasts in energy expenditure between sit‑and‑wait and actively foraging lizards.  Bulletin of the Ecological Society of America 61:94.

*Karasov, W. H. (1980) Water flux and water requirements of antelope ground squirrels in laboratory and field.  American Zoologist 20:742.

*Karasov, W. H., and J. M. Diamond (1981) Intestinal glucose transport varies with taxa and diet.  American Zoologist 21:1030.

Lehrer, P. H., and W. H. Karasov (1981) Energetic efficiency of foraging on varying resources in whiptail lizards (Cnemidophorus tigris).  American Zoologist 21:924.

*Karasov, W. H., and J. M. Diamond (1983) Effect of dietary carbohydrate on intestinal glucose transport in mice.  Physiologist 25:241.

*Karasov, W. H., D. H. Solberg, and J. M. Diamond (1983) Adaptation of intestinal nutrient transport in endotherms.  American Zoologist 23:489.

*Anderson, R. A., and W. H. Karasov (1983) Energetic implications of widely foraging predation in Cnemidophorus.  American Zoologist 23:978.

Solberg, D. H., W. H. Karasov, and J. M. Diamond (1985) Adaptation of intestinal nutrient transport to dietary protein levels.  Federation Proceedings 44:811.

Debnam, E. S., W. H. Karasov, and C. S. Thompson (1986) Hyperglycemia and glucose transport across the rat small intestine.  An in vitro and in vivo study.  Journal of Physiology 376:36P.

*Karasov, W. H., and E. S. Debnam (1986) Rapid adaptation of intestinal glucose absorption.  Federation Proceedings 43:537.

Debnam, E. S., W. H. Karasov, and C. S. Thompson (1986) Hyperglycemia:  evidence for sequential changes in glucose transport in the microvillus membrane and basolateral membrane of jejunal enterocytes.  Gut 27:A627‑A628.

*Meyer, M. W., and W. H. Karasov (1986) Creosote resin deters feeding by desert woodrats.  Abstracts of the American Society of Mammalogy 66th annual meeting, p.33.

*Karasov, W. H., and M. W. Meyer (1986) Water and protein imbalances in desert woodrats in wintertime.  American Zoologist 26:9A.

*Meyer, M. W., and W. H. Karasov (1986) Does creosote resin cause low protein digestibility in the desert woodrat?  American Zoologist 26:55A.

Meyer, M. W., and W. H. Karasov (1987) Physiological bases for tannic acid’s feeding deterrence in prairie voles.  Abstracts of the American Society of Mammalogy 67th annual meeting.

Munger, J. C., and W. H. Karasov (1987) Effects of Hymenolepis citelli infection on the energy budget of white-footed mice, Peromyscus leucopus.  Abstracts of the American Society of Parasitologists 62nd annual meeting.

*Karasov, W. H., B. W. Darken, and M. C. Bottum (1987) Tests for nonspecific and specific adaptation of intestinal choline transport.  Physiologist 30:191.

Levey, D. L., W. H. Karasov, and C. Martinez del Rio (1987) Switching diets from insects to fruits:  The digestive efficiencies of Robins and Starlings.  Bulletin of the Ecological Society of America 68(3):350.

Martinez del Rio, C., D. L. Levey, and W. H. Karasov (1987) Physiological basis and ecological correlates of sugar preferences in a North American frugivore, the cedar waxwing.  Bulletin of the Ecological Society of America 68(3):358.

Anderson, R. A., and W. H. Karasov (1988) Reproductive energetics of the wide-intensive foraging lizard, Cnemidophorus tigris.  Bulletin of the Ecological Society of America 69(2):57-58.

Meyer, M. W., and W. H. Karasov (1988) Do plant “digestibility-reducing defense compounds” reduce digestibility?  Bulletin of the Ecological Society of America 69(2):231.

Munger, J. C., and W. H. Karasov (1988) Sublethal parasites and host energy budgets:  Tapeworm infection in white-footed mice.  Bulletin of the Ecological Society of America 69(2):242.

*Karasov, W. H., and B. W. Darken (1989) Rebound scurvy:  Does reduced intestinal transport of ascorbic acid play a role?  Faseb Journal 3:A1154.

Meyer, M. W., and W. H. Karasov (1989) There is no simple dichotomy between digestibility-reduction and toxicity in action of plant secondary chemicals.  Bulletin of the Ecological Society 69(2):231.

Munger, J. C., and W. H. Karasov (1989) Sublethal parasites in white-footed mice:  Impact on survival and reproduction.  Bulletin of the Ecological Society 69(2):242.

*Karasov, W. H., and M. W. Meyer (1989) Foraging and physiology:  Incorporating physiological constraints into optimal foraging analyses.  Proceedings of the Fifth International Theriological Congress I:327.

*Karasov, W. H., and M. W. Meyer (1989) Digesta retention, nutrient absorption, and digestive efficiency in small herbivores.  Proceedings of the Fifth International Theriological Congress I:450.

*Karasov, W. H. (1989) Can terrestrial vertebrates digest and absorb chitin?  American Zoologist 29:24A.

Carey, C., and W. H. Karasov (1990) Effect of temperature on digestion of boreal toads, Bufo boreus boreus.  American Zoologist 33:34A.

Dykstra, C. R., and W. H. Karasov (1990) Adaptation of intestinal absorption to increased energy demand in House Wrens.  American Zoologist 30:66A.

Hume, I. D., and W. H. Karasov (1990) Diurnal and positional differences in acetate uptake in the hindgut of prairie voles.  Physiologist 33:A-34.

*Karasov, W. H. (1990) Dietary regulation of intestinal nutrient transport.  Physiologist 33:A-3.

Munger, J. C., and W. H. Karasov (1990) Do bot fly larvae affect metabolic rate and body temperature of small mammal hosts?  Bulletin of the Ecological Society of America 71(2):265.

Titus, E., W. H. Karasov, and G. A. Ahearn (1990) Effects of dietary composition on intestinal nutrient transport in euryhaline teleost.  Physiologist 33:A-34.

*Karasov, W. H., and C. R. Dykstra (1991) Paracellular absorption in normally fed insectivorous House Wrens.  FASEB Journal 5(5):A1137.

Dykstra, C. R., and W. H. Karasov* (1992) Test of energetic limitation to brood size in house wrens.  American Zoologist 32:91A.

Afik, D., and W. H. Karasov (1992) Dietary modulation of digestive efficiency in a seasonally frugivorous bird.  American Zoologist 32:99A.

Guglielmo, C. G., W. H. Karasov, and W. J. Jakubas (1993) Nutritional costs of a plant secondary metabolite to an avian herbivore.  Bulletin of the Ecological Society of America 74(2):260.

Jakubas, W. J., C. G. Guglielmo and W. H. Karasov (1993) Impact of plant secondary metabolites on energy and nutrient utilization in ruffed grouse.  Transactions of 55th Midwest Fish and Wildlife Conference, page 253.

*Karasov, W. H. (1994) Digestive mechanisms and constraints for fueling the metabolism of birds.  Physiologist 37:A-5.

Caviedes-Vidal, E., D. Afik, C. Martinez del Rio and W. H. Karasov (1994) Omnivory and dietary plasticity are not necessarily correlated:  Dietary modulation of intestinal enzymes.  Physiologist 37:A-81.

*Afik, D., E. Caviedes-Vidal and W. H. Karasov (1994) Passive absorption of glucose in three bird species:  Mediated glucose uptake alone cannot account for total glucose absorption.  Physiologist 37:A-81.

*Caviedes-Vidal, E., and W. H. Karasov (1995) Influences of diet composition on pancreatic enzyme activities in House Sparrows.  American Zoologist 35(5):78A.

*McWilliams, S. R., and W. H. Karasov (1995) Do variable reward feeding schedules affect digestive strategies of migratory birds?  American Zoologist 35(5):96A.

Lepczyk, C. A., and W. H. Karasov (1996).  The effect of ephemeral food restrictions on growth of House Sparrows (Passer domesticus).  Bulletin of the Ecological Society of America 77(3):260.

Hatch, K. A., S. R. McWilliams, W. H. Karasov, I.W. Treichel, M. E. Cook and W. P. Porter (1996).  The impact of metabolic and nutritional state on the use of tissue carbon isotope ratios to determine diet.  Bulletin of the Ecological Society of America 77(3):186.

Mangione, A.M., M.D. Dearing, and W.H. Karasov (1997) A test of the hypothesis of local adaptation: disparate tolerance of desert woodrats to creosote resin.  Bulletin of the Ecological Society of America 78(4):279.

Dearing, M.D., A.M. Mangione, and W.H. Karasov (1997) Detoxification of juniper terpenes by specialist and generalist woodrats.  Bulletin of the Ecological Society of America 78(4):78.

*Karasov, W.H. and B. Pinshow (1997) Body composition of spring migrant Blackcaps in the Negev Desert. American Zoologist 37(5):84A.

Ciminari, E., C. Codorniu, E. Caviedes-Vidal, S. McWilliams and W.H. Karasov (1998)  Dietary modulation of the intestinal disaccharidases maltase and sucrase in the Canada geese, Branta canadensis from North America. Biocell 22(3).

Ciminari, E., C. Codorniu, E. Caviedes-Vidal, S. McWilliams and W.H.Karasov (1998) Disaccharidase activity in the caeca of Canada geese, Branta canadensis from North America. Biocell. 22(3).

Caviedes Vidal, E. , E. Ciminari  and W. H. Karasov (1998) Ontogenetic variation of pancreatic enzymes in Passer domesticus. Biocell. 22(3).

Chediack J., Caviedes Vidal E., Karasov W. and Pestchanker M. (1998) Comparison of passive permeability of two sugars in Passer domesticus. BioCell.  Accepted.

Dearing, M.D., A.M. Mangione, and W.H. Karasov (1998) Diet breadth in mammalian herbivores: tests of the nutrient-constraints and detoxification-limitation hypotheses.  American Zoologist 38(5):14A.

Mangione, A.M., M.D. Dearing, and W.H. Karasov (1998) A plant secondary metabolite increases minimum water requirements of desert woodrats. American Zoologist 38(5):114A.

Chang, P., K. Castle, and W.H. Karasov (1998) Passive L-glucose absorption in mice in vitro and in vivo.  American Zoologist 38(5):121A.

Caviedes-Vidal, E., J.G. Chediack, W.H. Karasov, and M. Pestchanker. (1998). Passive absorption of D-mannitol and L-arabinose in Passer domesticus.  American Zoologist 38(5):115A.

Karasov, W.H. (1999). Optimal digestive responses to changing diet and foraging costs.  In: Adams, N. & Slotow, R. (Eds), Proc. 22 Int. Ornithol. Congr. Durban, University of Natal.

Sakaguchi, E., A. Mangione, F. Fournier, and W.H. Karasov (2000) Coprophagy and digesta transit in desert woodrats (Neotoma lepida). Proceedings of the Comparative Nutrition Society 3:187-193.

*Caviedes-Vidal, E., W. H. Karasov.  (2000) Developmental changes in digestive enzymes of nestling house sparrows, Passer domesticus. Amercian Zoologist 40:968

Chang, M. H., Chediack, J. G., Caviedes-Vidal, E., and Karasov, W. H. (2000) Enhanced passive absorption in the presence of luminal nutrients in house sparrows. American Zoologist 40:970

Edblom, T. L., and Karasov, W. H. (2000) Solar ultraviolet-B radiation does not affect hatching success of Rana pipiens (northern leopard frog) eggs under natural conditions along a contamination gradient. American Zoologist 40:1006

Fournier, F., Karasov, W. H., and Meyer, M. W. (2000) The effect of feather development on the pharmacokinetics and oral bioavailability of methylmercury in common loon chicks.  American Zoologist 40:1018

Morris, K. A., Karasov, W. H., and Caviedes-Vidal, E. (2000) Does digestive physiology limit re-fueling rates in migratory yellow-rumped warblers (Dendroica coronata)? American Zoologist 40:1139

Chang, M. H. and Karasov, W. H. Similar extent and rate of absorption of D- and L-glucose in house sparrows, Passer domesticus. FASEB Journal 15(5), A830. 2001.

McWilliams, S.R., S.B. Kearney, and W.H. Karasov (2001) Diet preferences of warblers for specific fatty acids in relation to nutritional requirements and digestive capabilities.  American Zoologist 41(6):1523.

Fasulo, V., E. Caviedes-Vidal, and W.H. Karasov (2001) In vivo modulation of intestinal paracellular absorption by mediated glucose transport.  American Zoologists 41(6):1440.

*Karasov, W.H., E.L. McSwan, and J. Wright (2001).  Begging and digestion by nestlings Pied flycatchers in relation to short- and long-term nutritional need.  American Zoologist 41(6):1490.

*Chediack, J.G., E. Caviedes-Vidal, and W.H. Karasov (2001)  Intestinal passive absorption of hydrosoluble compounds is size dependent and modulated by luminal nutrients in house sparrows (Passer domesticus).  American Zoologist 41(6):1410.

McWilliams, S. R. and W. H. Karasov (2003)  Spare capacity in the digestive system of a migratory songbird and its ecological significance. Abstracts of SICB Annual Meeting 248.

Skopec, M. M. and W. H. Karasov (2003)  Do salivary proline-rich proteins decrease the metabolic cost of tannin consumption? Abstracts of SICB Annual Meeting 308.

*Green, A.K., M.M. Skopec, and W.H. Karasov (2003) Flavonoids decrease 3-O-methyl D-glucose absorption in American Robins.  Integrative and Comparative Biology 43:1007.

Green, A.K., S.L. Haley, W.H. Karasov, D.M. Barnes, and D.M. Dearing (2003)  Total capacity of P-glycoprotein higher in the specialist herbivore Neotoma stephensi than the congeneric generalist N. albigula.  Integrative and Comparative Biology 43:1008.

McWhorter, T.J., W.H. Karasov and A.K. Green. (2004) How the American robin absorbs glucose. Integrative and Comparative Biology 44(6): 602.

Lavin, S.R., T.J. McWhorter and W.H. Karasov. (2004) Do birds exhibit greater paracellular absorption than mammals?  Integrative and Comparative Biology 44(6): 717.

McWhorter, T.J., W.H. Karasov and A.K. Green. (2005)  How the American robin absorbs glucose.  FASEB Journal 19(4): A752-A752 Part 1 Suppl. S, MAR 4 2005. [Featured topic presented paper and poster, XXXV International Congress of Physiological Sciences, San Diego, CA. March 31- April 6, 2005.]

*Caviedes-Vidal, E., J.G. Chediack, A.P. Cruz-Neto, and W.H. Karasov (2004)  Sugar absorption in bats.  Are they mammals or birds? Integrative and Comparative Biology 44(6): 534.

Fassbinder-Orth, C. A. & Karasov, W. H. (2005) Are gut immune and digestive functions regulated independently? Integrative and Comparative Biology 45(6), 1130.

Lavin, S. R., McWhorter, T. J. & Karasov, W. H. (2005) Differences in paracellular absorption: a function of solvent drag or pore size? Integrative and Comparative Biology 45(6), 1030.

McWhorter, T. J., Hartman Bakken, B., Karasov, W. H. & Martínez del Rio, C. (2005) Hummingbirds fuel high metabolism with both paracellular and carrier-mediated intestinal glucose absorption. Integrative and Comparative Biology 45(6), 1043.

*Tracy, C. R., McWhorter, T. J., Korine, C., Pinshow, B. & Karasov, W. H. (2005) A mystery solved: how the Egyptian fruit bat absorbs sugar. Integrative and Comparative Biology 45(6), 1085.

Tracy, C. R., McWhorter, T. J., Wojciechowski, M. S., Korine, C., Karasov, W. H. & Pinshow, B. 2005 Relatively low paracellular absorption in a reptile, the Egyptian mastigure, Uromastyx aegyptius. Integrative and Comparative Biology 45(6), 1203.

Tracy, C. R., Wojciechowski, M. S., McWhorter, T. J., Yosef, R., Karasov, W. H. & Pinshow, B. (2005) Carbohydrate absorption in migrating blackcaps, upon arrival and after refeeding, at a stopover in southern Israel. Integrative and Comparative Biology 45(6), 1085.

*Karasov, W.H., McWhorter, T.J., Caviedes-Vidal, E. (2006) Phylogenetic and body size patterns in intestinal paracellular solute absorption. FASEB Journal 20(5):A1275 Part 2.

*McWhorter, T.J. and W.H. Karasov.  (2006)  Paracellular absorption varies with body size in primates. FASEB Journal 20(5):A1275 Part 2

Wojciechowski M.S., Tracy C.R., McWhorter T.J., Yosef R., Karasov W.H., Pinshow B. (2006) Blackcaps at migratory stopovers compensate for reduced gut with increased paracellular nutrient uptake. J. Ornithol. 147 suppl. 1:274

Caviedes-Vidal, E. and W.H. Karasov.  (2007)  Digestive plasticity in birds: Mechanisms and ecological consequences. Comparative Biochemistry and Physiology, Part A 148:S132.

*Karasov, W.H. and E. Caviedes-Vidal.  (2007)  The intestine as a site of absorption, elimination, and action of toxins in vertebrates. Comparative Biochemistry and Physiology, Part A 148:S132.

Naya, D.E., W.H. Karasov and F. Bozinovic.  (2007) Gut size flexibility in laboratory mice and rats: A meta-analysis. Comparative Biochemistry and Physiology, Part A 148:S133.

Brzek, P., E. Caviedes-Vidal, and W.H. Karasov (2007) Glucose absorption in House sparrow nestlings does not depend on age or diet. Integrative and Comparative Biology (http://icb.oxfordjournals.org/cgi/reprint/47/suppl_1/e1)

Kohl, K., P. Brzek, E. Caviedes-Vidal, and W.H. Karasov (2007) Diet-induced plasticity in intestinal maltase activity in nestling house sparrows during development. Integrative and Comparative Biology (http://icb.oxfordjournals.org/cgi/reprint/47/suppl_1/e1)

Karasov, W.H. (2008) Digestive features of birds confer plastic responses to changing food intake and composition. Comparative Biochemistry and Physiology 150A (3/Supplement): S61.

Brzek, P., E. Caviedes-Vidal, W.H. Karasov (2009) Paracellular and total glucose absorption increase with age in nestling House sparrows. Integrative and Comparative Biology 49:e21.

*Brzek, P., K. Kohl, E. Caviedes-Vidal, and W.H. Karasov (2009) Activity of intestinal carbohydrases responds to multiple dietary signals in nestling House sparrows. Integrative and Comparative Biology 49:e22.

Kohl, K., P. Brzek, E. Caviedes-Vidal, and W.H. Karasov (2009) Phenotypic plasticity of intestinal dissacharidase activity is fully reversible in young House sparrows. Integrative and Comparative Biology 49:e93.

Baston, J.I., J.G. Chediack, F.D. Cid, W.H. Karasov, and E. Caviedes-Vidal (2009) Daily expression modulation of SGLT1 in rat jejunum during ontogeny. Integrative and Comparative Biology 49:e197.

Gatica Sosa, C, J.G. Chediack, M. Juri Ayub, W.H. Karasov, and E. Caviedes-Vidal (2009) Expression of the intestinal brush border disaccharidases and Na+/glucose cotransporter genes of zebra finch (Taeniopygia guttata). Integrative and Comparative Biology 49:e233.

Zhang, Z. Q., M. E. Arce, G. M. Ciuffo, W. H. Karasov, and E. Caviedes-Vidal. 2010. Intestinal morphometrics in flying and non-flying mammals. Integrative and Comparative Biology 50:E318-318.

Kohl, K. D., P. Brzek, E. Caviedes-Vidal, and W. H. Karasov. 2010. Matching between dietary preferences and digestive capacity in passerine birds. Integrative and Comparative Biology 50:E92-E92.

Killpack, T., N. Singh, and W. H. Karasov. 2010. Effect of chronic food restriction on gut morphology and digestive enzymes in nestling House sparrows. Integrative and Comparative Biology 50:E90-E90.

Brzek, P., E. Caviedes-Vidal, and W. H. Karasov. 2010. House sparrow fledglings leave the nest digestively immature but more flexible than adults. Integrative and Comparative Biology 50:E19-E19.

Cary, T. L., M. E. Ortiz-Santaliestra, N. D. Schmidt, and W. H. Karasov. 2011. Assessment of development and immune status of northern leopard frogs (Rana pipiens) following polybrominated diphenyl ether (PBDE) exposure. Integrative and Comparative Biology 51:E172-E172.

Killpack, T. L., K. Stone, and W. H. Karasov. 2011. Body composition and developmental maturity in chronically restricted nestling House sparrows. Integrative and Comparative Biology 51:E213-E213.

Liu, J. N. and W. H. Karasov. 2011. Energetics, thermoregulation, and energy expenditure during winter in the subtropical hibernating bat Hipposideros terasensis. Integrative and Comparative Biology 51:E82-E82.

Anderson, R. A., W. H. Karasov, and K. A. Nagy. 2012. Ecological cost of growth in a free-ranging lizard. Integrative and Comparative Biology 52:E6-E6.

Cary, T. L. and W. H. Karasov. 2012. Immunotoxicity may be a more sensitive endpoint for sublethal polychlorinated biphenyl exposure in the northern leopard frog. Integrative and Comparative Biology 52:E26-E26.

Killpack, T. L. and W. H. Karasov. 2012. Post-hatching ontogeny of adaptive antibody response to two distinct antigens in captive zebra finches. Integrative and Comparative Biology 52:E93-E93.

Killpack, T. L., Y. Oguchi, and W. H. Karasov. 2012. Ontogeny of constitutive innate immune function in free-living altricial house sparrow nestlings. Integrative and Comparative Biology 52:E275-E275.

Liu, J. and W. H. Karasov. 2012. Energetics and use of torpor during summer in a subtropical bat, the Formosan leaf-nosed bat Hipposideros terasensis. Integrative and Comparative Biology 52:E107-E107.

Nan Tie, D. E., T. L. Killpack, and W. H. Karasov. 2012. Effect of food restriction and realimentation on growth, immune response, and body composition in Zebra finches. Integrative and Comparative Biology 52:E128-E128.

Price, E. R., L. Ruff, A. Guerra, and W. H. Karasov. 2012. Mice increase reliance on paracellular intestinal absorption in response to increased energy demand. Integrative and Comparative Biology 52:E141-E141.

Karasov, W. H., E. R. Price, L. Ruff, A. Guerra. 2012. Paracellular intestinal absorption increases in mice exposed to cold. FASEB 26:APR.

Cary, T. L., J. D. Pask, L. A. Rollns-Smith, W. H. Karasov. 2013. PCB-126 exposure lowered antimicrobial peptide secretion in juvenile northern leopard frogs. Integrative and Comparative Biology 53:E259-E259.

*Karasov, W. H. Research paradigms in nutritional ecology inspired by Ken Nagy. 2013. Integrative and Comparative Biology 53:E106-E106.

Killpack, T. L., E. Carrel, W. H. Karasov. 2013. Impact of food restriction on immune function in altricial house sparrow nestlings. Integrative and Comparative Biology 53:E110-E110.

Price, E. R., A. Brun, V. Fasulo, W. H. Karasov, E. Caviedes-Vidal. 2013. Paracellular absorption of nutrients in bats is high during intestinal luminal perfusions. Integrative and Comparative Biology 53: E173-E173.

Richman, S. E. Sr. R. McWilliams, J. O. Leafloor, W. H. Karasov. 2013. Growing Fast and Dying Young: Influence of Forage Quality on Growth and Survival of Arctic Avian Herbivores. Integrative and Comparative Biology 53:E180-E180.

Samuni-Blank, M., I. Izhaki, M. D. Dearing, W. H. Karasov, Y. Gerchman, K. Kohl, P. Lymberakis, P. Kurnath, Z. Arad. 2013. Divergent Behavioral Strategies in Three Congeneric Rodents for Dealing with Fruit Toxins. Integrative and Comparative Biology 53:E188-E188.

*Karasov, W.H., A. Brun, E.R. Price, M.N. Gontero-Fourcade, G. Fernandez-Marinone, A.P. Cruz Neto and E. Caviedes-Vidal (2014). Intestinal paracellular permeability to nutrients is higher in frugivorous bats than rodents. Integrative and Comparative Biology 54:E108

Price, E.R., E. Caviedes-Vidal, and W.H. Karasov (2014) Mechanistic correlates of paracellular absorption in protein specialists. Integrative and Comparative Biology 54:E169.

Price, E.R., A. Brun, E. Caviedes-Vidal, and W.H. Karasov (2015) Bats and birds share digestive adaptations to an aerial lifestyle. The Physiologist 58(1):29.

Price, E.R., K.H. Rott, E. Caviedes-Vidal, and W.H. Karasov (2015) An approach for comparing qPCR data across species, applied to paracellular nutrient absorption. Integrative and Comparative Biology 55 (Supplement 1):E146.

Price, E.R., A. Brun, E. Caviedes-Vidal, and W.H. Karasov (2015) Bats and birds share digestive adaptations to an aerial lifestyle. Integrative and Comparative Biology 55 (Supplement 1):E146.

Tsai, C.A., J.M. Yahn, and W.H. Karasov (2015) Warmer temperature increases toxicokinetic elimination of PCB and PBDE in larvae. Integrative and Comparative Biology 55 (Supplement 1):E344.

Cornelius, E.A., L. Regimbald, O. Love, F. Vezina and W.H. Karasov (2016) Immune function and body composition of a wild bird faced with food unpredictability. Integrative and Comparative Biology 56 (Supplement 1) E42

Rott, K.H., E. Caviedes-Vidal, and W.H. Karasov (2016) Intestinal enzyme activity in nestling house sparrows (Passer domesticus) not depressed by high dietary lipid content. Integrative and Comparative Biology 56 (Supplement 1) E361

Rott, K.H., E. Caviedes-Vidal, and W.H. Karasov (2016) Intestinal enzyme activity in nestling house sparrows (Passer domesticus) is modulated within 24 hours of a diet switch. Integrative and Comparative Biology 56 (Supplement 1) E189

Cornelius Ruhs, E, F. Vezina, and W.H. Karasov (2017) Do differing levels of food supplementation alter body composition and immune function in a wild residential bird?  Integrative and Comparative Biology 57 (Supplement 1) E230.

Latimer, C.E., S.J. Cooper, W.H. Karasov et al. (2017) Metabolic constraints differentially affect foraging and survival of birds in human-modified landscapes.  Integrative and Comparative Biology 57 (Supplement 1) E322.

Rott, K.H., E. Caviedes-Vidal, and W.H.Karasov (2017) Activity of intestinal enzymes responds to multiple dietary signals in birds. Integrative and Comparative Biology 57 (Supplement 1) E391.

Wang, L. and W. H. Karasov (2017) Diet composition modulates intestinal hydrolytic enzymes in white-footed mice (Peromyscus leucopus) FASEB Journal 31 (Supplement 1):889.4

Gatica-Sosa, C., P. Brzek, M.E. Magallanes, et al. (2018) Intestinal alpha-glucosidase transcriptional responses during ontogeny and diet adjustment in altricial birds. Integrative and Comparative Biology 58 (Supplement 1) E73

Kohl, K.D., A. Brun, E. Caviedes-Vidal, and W.H. Karasov (2018) Gut microbial ecology of nestling House Sparrows (Passer domesticus). Integrative and Comparative Biology 58 (Supplement 1) E122.

Brun, A., D. Mendez-Aranda, M.E. Magallanes, W.H. Karasov, C. Martinez del Rio, M. Baldwin, and E. Caviedes-Vidal (2019)  Evolution of intestinal α-glucosidases in vertebrates: Genomic and proteomic data upend previous hypotheses. Integrative and Comparative Biology (Supplement 1): in press

Brun, A. G.A. Barrett-Wilt, W.H. Karasov, and E. Caviedes-Vidal (2019) Proteomics of the enzyme proteins in the intestinal brush border membrane of vertebrates. Integrative and Comparative Biology (Supplement 1): in press.

Karasov, W.H., A. Brun, M.E. Magallanes, K.H. Rott, C. Ross, and E. Caviedes-Vidal (2019) Intestinal hydrolase transcriptional responses during rapid diet adjustment in nestling house sparrows (Passer domesticus). Physiologist 62(1): Supplement for Intersociety Meeting, Comparative Physiology: pg. 42